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Cicadas (Hemiptera: Cicadidae) from Panay, Philippines, with a new species and a new genus
Journal of Asia-Pacific Entomology 12 (2009) 293–295
Contents lists available at ScienceDirect
Journal of Asia-Pacific Entomology j o u r n a l h o m e p a g e : w w w. e l s e v i e r. c o m / l o c a t e / j a p e
Cicadas (Hemiptera: Cicadidae) from Panay, Philippines, with a new species and a new genus Young June Lee ⁎ Department of Ecology and Evolutionary Biology, University of Connecticut, 75 North Eagleville Road, Storrs, CT 06269, USA
a r t i c l e
i n f o
Article history: Received 19 April 2009 Revised 22 June 2009 Accepted 25 June 2009 Keywords: Checklist Ggomapsalta Platypleura lourensi Dundubia vaginata Ggomapsalta vernalis
a b s t r a c t This paper provides the first faunal checklist for Cicadidae (Hemiptera) from Panay, Philippines, comprising five species belonging to five genera. A new species, Platypleura lourensi sp. nov., and a new genus, Ggomapsalta gen. nov., are described. Dundubia vaginata (Fabricius, 1787) and Ggomapsalta vernalis (Distant, 1916) are newly recorded from Panay. Information on geographic distributions is provided. © Korean Society of Applied Entomology, Taiwan Entomological Society and Malaysian Plant Protection Society, 2009 Published by Elsevier B.V. All rights reserved.
Introduction
Taxonomic checklist
This paper is aimed for a contribution to the knowledge of cicada diversity in Panay, Philippines, providing its first faunal checklist. Panay is the fourth largest island of the Philippines. The previous records of the cicadas from Panay were made only by Hayashi (1987) and Duffels and Zaidi (2000) to record Cryptotympana socialis Hayashi, 1987 and Orientopsaltria fuliginosa (Walker, 1850). By describing a new species and a new genus and adding two species to the fauna in this paper, the cicada fauna of Panay now comprises five species belonging to five genera as listed below. This list of species is based on the literature that specifically mentioned the fauna of Panay Island and the specimens from the Institut royal des Sciences naturelles de Belgique, Brussels (IRSNB). The synonymic list for each species includes only the references that specifically mentioned the fauna of Panay, in addition to the original descriptions with the type localities (TL), which is listed as given in the original publications. Nomenclature for family, subfamily, and tribal classification follows that of Lee (2008), and morphological terminology follows that of Moulds (2005). Morphological measurements were made with a Mitutoyo™ vernier caliper in mm.
Family Cicadidae Subfamily Cicadinae Tribe Platypleurini Genus Platypleura Amyot and Audinet-Serville, 1843 Type species: Cicada stridula Linnaeus, 1758. Platypleura Amyot and Audinet-Serville, 1843: 465.
⁎ Fax: +1 860 465 9730. E-mail address: [email protected].
Platypleura lourensi sp. nov. (Figs. 1, 2) Type material. Holotype: male (Fig. 1), Panay, Aklan, 1 km N. of Logtugan, 12 km W. of Libacao, sec. regrown forest, 105 m, 11°24.691′N 122°18.528′E, 2 IX 2008, J.H. Lourens (IRSNB). Paratypes: 1 male, 17 females, same data as holotype (IRSNB). Etymology. The species is named for the collector of a large collection of the cicadas from the Philippines, including this new species, Mr. J.H. Lourens, who made this paper possible. Measurements of types (2 males, 17 females). Length of body: male 21.8 (21.5–22.0), female 23.8 (22.6–25.2); width of head including eyes: male 10.5 (10.4–10.6), female 11.2 (10.7–11.7); wing span: male 68.4 (68.1–68.7), female 74.7 (71.5–78.4). Diagnosis. This species resembles Hamza ciliaris (Linnaeus, 1758) in the coloration of the fore and the hind wings but is distinguished by lack of fuscous fasciae on the bases of the radial cell and the medial cell as well as along the cubital cell 2 and the anal cell 1 of the hind wing. In male specimens, this species is easily distinguished by a nearly complete timbal cover against the incomplete, uniquely shaped timbal cover of Hamza ciliaris. This species is also allied to Platypleura hilpa
1226-8615/$ – see front matter © Korean Society of Applied Entomology, Taiwan Entomological Society and Malaysian Plant Protection Society, 2009 Published by Elsevier B.V. All rights reserved. doi:10.1016/j.aspen.2009.06.007
294
Y.J. Lee / Journal of Asia-Pacific Entomology 12 (2009) 293–295
Fig. 1. Platypleura lourensi sp. nov., holotype, male, Aklan, Panay (IRSNB). (A) dorsal view. (B) ventral view.
Walker, 1850 but is distinguished by a considerably developed lateral pronotal collar and the fuscous outer margin of the anal cell 2 (vannus). This species is different from any other species of Platypleura in having the timbal cover of which inner corner is continuously rounded (not angled) to the anterior margin of the abdominal tergite 2. Description of male (Fig. 1). Head light green to greenish ochraceous with the following black marks: a median large spot enclosing ocelli; two pairs of longitudinally arranged small spots on the sides of the median spot, of which both of each pair are nearly connected to each other; a pair of oblique marks between the anterior one of the longitudinally arranged small spots and compound eyes; and a pair of small marks or fasciae on supra antennal plates. Distance between lateral ocelli and compound eyes slightly wider than twice the distance between two lateral ocelli. Postclypeus not swollen. Antenna mostly dark brown except fuscous pedicel. Ventral part of head light green to greenish ochraceous with black marks and densely covered with silvery hairs and white pollinosity. Postclypeus with transverse fasciae along transverse grooves, of which medial ends (or paramedical ends in case of anterior fasciae) are continued to each other. Anteclypeus with a pair of large marks, which occupy most part of anteclypeus except a narrow longitudinal median part. Rostrum black apically; slightly passing posterior margin of hind coxae. Lorum with a patch on posterior part. Gena with a broad transverse fascia between antenna and compound eye. Pronotum light green to greenish ochraceous. Inner area of pronotum with indistinct narrow brown patches on paramedian and
lateral fissures. Pronotal collar with a pair of transversely arranged medial tiny spots on anterior margin and dark brown patches on lateral submargin. Lateral pronotal collar broadly developed and angulated with an angle of about 100°. Mesonotum green to greenish ochraceous with the following marks: a median longitudinal black fascia, which is broadest at about center of mesonotum and sometimes elongated posteriad to touch anterior margin of cruciform elevation; a pair of small roundish black spots enclosing scutal depressions; a pair of obconical large black spots on submedian sigilla; and a pair of longitudinal long triangular castaneous patches on lateral sigilla, of which anterior ends reach anterior margin of mesonotum. Cruciform elevation ochraceous with delicate transverse fascia along anterior margin. Ventral part of thorax light green to greenish ochraceous with black marks and densely covered with silvery hairs and white pollinosity. Legs ochraceous to greenish ochraceous. Tarsi and pretarsal claws with darker parts. Fore wing venation green basally and reddish brown apically. Fore wing entirely furnished with gray and brownish cloudy marks. Hind wing ochraceous with a fuscous band along outer margin and a large fuscous spot covering over about apical halves of costal, radial, and medial cells and basal areas of apical cells 1, 2, 3, 4, and 5, and sometimes 6, which is continued with the band anteriorly. Operculum greenish ochraceous; semicircular, transversely wide; short, reaching posterior margin of sternite II. Two opercula slightly or not overlapping medially. Abdomen obconical, short, shorter than distance from head to cruciform elevation; black with greenish ochraceous caudal margin on each of tergites 2–7. Tergite 8 entirely black. Timbal cover greenish ochraceous to ochraceous; semicircular, wider than long. Timbal slightly exposed paramedially in dorsal view. Inner corner of timbal cover continuously rounded to anterior margin of tergite 2. Ventral part of abdomen dark ochraceous. Male genitalia (Fig. 2): Pygofer ellipsoidal with acuminate apex (wide dorsal beak), but rather triangular at about upper half, in ventral view. Uncus not bifurcate, thick in lateral view, with apex roundly expanded downward. Aedeagus not tapering down to apex. Upper and basal lobes of pygofer not prominent. Distribution. Philippines (Panay). Tribe Cryptotympanini Genus Cryptotympana Stål, 1861 Type species: Tettigonia atrata Fabricius, 1775. Cryptotympana Stål, 1861: 613. Cryptotympana socialis Hayashi, 1987 Cryptotympana socialis Hayashi, 1987: 13 [TL: Antique, Isl. Panay]. Distribution. Philippines (Panay, Cebu, and Negros). Tribe Cicadini Subtribe Dundubiina Genus Orientopsaltria Kato, 1944 Type species: Dundubia duarum Walker, 1857. Orientopsaltria Kato, 1944: 6. Orientopsaltria fuliginosa (Walker, 1850) Dundubia fuliginosa Walker, 1850: 60 [TL: Philippine Islands]. Orientopsaltria fuliginosa: Duffels and Zaidi, 2000: 275.
Fig. 2. Male pygofer of Platypleura lourensi sp. nov. (A) ventral view. (B) lateral view.
Material examined. 1 male, 2 females, Panay, Iloilo, Maasin, riverbed forest, replant, 100 m, 10°53.882′N 122°24.595′E, 29 VIII 2008, J.H. Lourens (IRSNB); 5 males, 9 females, Panay, Iloilo, E. of Mt. Baloy, Bgy Supanga, 32 km W. of Calinog, 570 m, 11°09.460′N 122°21.279′E, 30–31 VIII 2008, J.H. Lourens (IRSNB). Distribution. Philippines (on most Philippine islands including Panay).
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developed. Fore femur with primary, secondary, and two small subapical spines. Wings hyaline without infuscation. Operculum short and wide, not reaching posterior margin of sternite II. Abdomen slightly longer than distance from head to cruciform elevation. Timbal cover absent. Pygofer nearly diamond-shaped in ventral view. Upper lobe of pygofer absent. Dorsal beak dull, not acute, triangular. Median lobe of uncus triangular, slender, and very long. A pair of clasper-like processes protruding downward. Etymology. The generic name is a feminine noun derived from the Korean ggoma- and the Latin psalta, meaning “a small musician”. Ggomapsalta vernalis (Distant, 1916) (Fig. 3) Pauropsalta vernalis Distant, 1916: 204 [TL: Mt. Makiling, Luzon, Philippine Islands]. Material examined. 1 male, Panay, Iloilo, E. of Mt. Baloy, Bgy Supanga, 32 km W. of Calinog, 570 m, 11°09.460'N 122°21.279′E, 30–31 VIII 2008, J.H. Lourens (IRSNB). Distribution. Philippines (Luzon and Panay). Remarks. This species is newly recorded from Panay. Acknowledgements
Genus Dundubia Amyot and Audinet-Serville, 1843 Type species: Tettigonia vaginata Fabricius, 1787. Dundubia Amyot and Audinet-Serville, 1843: 470.
I am indebted to Dr. Jerome Constant (Institut royal des Sciences naturelles de Belgique, Brussels) for the loan of specimens. I am grateful to anonymous reviewers for suggestions that improved the manuscript. This work benefited from support from the University of Connecticut and the National Science Foundation under Grant Numbers DEB 05-29679 and DEB 07-20664. Any opinions, findings, and conclusions or recommendations expressed in this material are those of the author and do not necessarily reflect the views of the NSF.
Dundubia vaginata (Fabricius, 1787) Tettigonia vaginata Fabricius, 1787: 266 [TL: Sumatra].
References
Fig. 3. Ggomapsalta vernalis (Distant, 1916), male, Iloilo, Panay (IRSNB). (A) dorsal view. (B) ventral view. (C) ventral view of pygofer. (D) lateral view of pygofer.
Material examined. 3 males, 1 female, Panay, Iloilo, E. of Mt. Baloy, Bgy Supanga, 32 km W. of Calinog, 570 m, 11°09.460′N 122°21.279′E, 30–31 VIII 2008, J.H. Lourens (IRSNB). Distribution. S. China, Philippines (on most Philippine islands including Panay), Malay Archipelago, S. Myanmar, and India. Remarks. This species is newly recorded from Panay. Subfamily Cicadettinae Tribe Cicadettini Genus Ggomapsalta gen. nov. Type species: Pauropsalta vernalis Distant, 1916. Diagnosis. This genus is similar to Pauropsalta Goding and Froggatt, 1904 in having the following common features: a wide head, which is wider than the base of the mesonotum; hyaline wings, which have no infuscation; and the hind wing, which has only five apical cells (Figs. 3A, B). But this genus is distinguished by the nearly unicolor body and the unique structure of the male genitalia (Figs. 3C, D): upper lobe of pygofer is absent; dorsal beak is dull, not acute, triangular; median lobe of uncus is triangular, slender, and very long; a pair of clasper-like processes protrude downward. Description. Body tiny, about 12 mm long. Head including eyes wider than base of mesonotum. Distance between lateral ocelli and compound eyes slightly wider than distance between two lateral ocelli. Postclypeus scarcely swollen. Anterolateral pronotal collar not
Amyot, C.J.B., Audinet-Serville, J.G., 1843. Homoptères. Homoptera Latr. In: Histoire naturelle des insectes, hémiptères. Paris: Librairie Encyclopédique de Roret. pp. 455–588. Distant, W.L., 1916. On the Cicadidae found in the Philippine Islands. Entomologist 49, 203–204. Duffels, J.P., Zaidi, M.I., 2000. A revision of the cicada genus Orientopsaltria Kato (Homoptera, Cicadidae) from Southeast Asia. Tijdschrift voor Entomologie 142, 195–297, 6 pls. Fabricius, J.C., 1787. Ryngota. p. 260–275 In: Mantissa insectorum sistens species nuper detectas adiectis synonymis, observationibus, descriptionibus, emendationibus, 2. pp. 1–382. Goding, F.W., Froggatt, W.W., 1904. Monograph of the Australian Cicadidae. Proc. Lin. Soc. N. S. W. 29, 561–668, pls. 18–19. Hayashi, M., 1987. A revision of the genus Cryptotympana (Homoptera, Cicadidae). Part II. Bulletin of the Kitakyushu Museum of Natural History 7, 1–109. Kato, M., 1944. Cicadidae from Sumatra and Malaya presented by Mr. T. Araki. Bulletin of the Cicadidae Museum, Tokyo 14, 1–8 (In Japanese). Lee, Y.J., 2008. A checklist of Cicadidae (Insecta: Hemiptera) from Vietnam, with some taxonomic remarks. Zootaxa 1787, 1–27. Linnaeus, C., 1758. Systema naturae, per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio decima, reformata, 1. Holmiae 1–824. Moulds, M.S., 2005. An appraisal of the higher classification of cicadas (Hemiptera: Cicadoidea) with special reference to the Australian fauna. Rec. Aust. Mus. 57, 375–446. Stål, C., 1861. Genera nonnulla nova cicadinorum. Annales de la Société entomologique de France (1) 4, 613–622. Walker, F., 1850. List of the specimens of homopterous insects in the collection of the British Museum, Part 1. London, pp. 1–260, pls. 1–2. Walker, F., 1857. Catalogue of the homopterous insects collected at Sarawak, Borneo, by Mr. A. R. Wallace, with descriptions of new species. Journal of the Proceedings of the Linnean Society, Zoology 1, 141–175, pls. 7–8.
Contents lists available at ScienceDirect
Journal of Asia-Pacific Entomology j o u r n a l h o m e p a g e : w w w. e l s e v i e r. c o m / l o c a t e / j a p e
Cicadas (Hemiptera: Cicadidae) from Panay, Philippines, with a new species and a new genus Young June Lee ⁎ Department of Ecology and Evolutionary Biology, University of Connecticut, 75 North Eagleville Road, Storrs, CT 06269, USA
a r t i c l e
i n f o
Article history: Received 19 April 2009 Revised 22 June 2009 Accepted 25 June 2009 Keywords: Checklist Ggomapsalta Platypleura lourensi Dundubia vaginata Ggomapsalta vernalis
a b s t r a c t This paper provides the first faunal checklist for Cicadidae (Hemiptera) from Panay, Philippines, comprising five species belonging to five genera. A new species, Platypleura lourensi sp. nov., and a new genus, Ggomapsalta gen. nov., are described. Dundubia vaginata (Fabricius, 1787) and Ggomapsalta vernalis (Distant, 1916) are newly recorded from Panay. Information on geographic distributions is provided. © Korean Society of Applied Entomology, Taiwan Entomological Society and Malaysian Plant Protection Society, 2009 Published by Elsevier B.V. All rights reserved.
Introduction
Taxonomic checklist
This paper is aimed for a contribution to the knowledge of cicada diversity in Panay, Philippines, providing its first faunal checklist. Panay is the fourth largest island of the Philippines. The previous records of the cicadas from Panay were made only by Hayashi (1987) and Duffels and Zaidi (2000) to record Cryptotympana socialis Hayashi, 1987 and Orientopsaltria fuliginosa (Walker, 1850). By describing a new species and a new genus and adding two species to the fauna in this paper, the cicada fauna of Panay now comprises five species belonging to five genera as listed below. This list of species is based on the literature that specifically mentioned the fauna of Panay Island and the specimens from the Institut royal des Sciences naturelles de Belgique, Brussels (IRSNB). The synonymic list for each species includes only the references that specifically mentioned the fauna of Panay, in addition to the original descriptions with the type localities (TL), which is listed as given in the original publications. Nomenclature for family, subfamily, and tribal classification follows that of Lee (2008), and morphological terminology follows that of Moulds (2005). Morphological measurements were made with a Mitutoyo™ vernier caliper in mm.
Family Cicadidae Subfamily Cicadinae Tribe Platypleurini Genus Platypleura Amyot and Audinet-Serville, 1843 Type species: Cicada stridula Linnaeus, 1758. Platypleura Amyot and Audinet-Serville, 1843: 465.
⁎ Fax: +1 860 465 9730. E-mail address: [email protected].
Platypleura lourensi sp. nov. (Figs. 1, 2) Type material. Holotype: male (Fig. 1), Panay, Aklan, 1 km N. of Logtugan, 12 km W. of Libacao, sec. regrown forest, 105 m, 11°24.691′N 122°18.528′E, 2 IX 2008, J.H. Lourens (IRSNB). Paratypes: 1 male, 17 females, same data as holotype (IRSNB). Etymology. The species is named for the collector of a large collection of the cicadas from the Philippines, including this new species, Mr. J.H. Lourens, who made this paper possible. Measurements of types (2 males, 17 females). Length of body: male 21.8 (21.5–22.0), female 23.8 (22.6–25.2); width of head including eyes: male 10.5 (10.4–10.6), female 11.2 (10.7–11.7); wing span: male 68.4 (68.1–68.7), female 74.7 (71.5–78.4). Diagnosis. This species resembles Hamza ciliaris (Linnaeus, 1758) in the coloration of the fore and the hind wings but is distinguished by lack of fuscous fasciae on the bases of the radial cell and the medial cell as well as along the cubital cell 2 and the anal cell 1 of the hind wing. In male specimens, this species is easily distinguished by a nearly complete timbal cover against the incomplete, uniquely shaped timbal cover of Hamza ciliaris. This species is also allied to Platypleura hilpa
1226-8615/$ – see front matter © Korean Society of Applied Entomology, Taiwan Entomological Society and Malaysian Plant Protection Society, 2009 Published by Elsevier B.V. All rights reserved. doi:10.1016/j.aspen.2009.06.007
294
Y.J. Lee / Journal of Asia-Pacific Entomology 12 (2009) 293–295
Fig. 1. Platypleura lourensi sp. nov., holotype, male, Aklan, Panay (IRSNB). (A) dorsal view. (B) ventral view.
Walker, 1850 but is distinguished by a considerably developed lateral pronotal collar and the fuscous outer margin of the anal cell 2 (vannus). This species is different from any other species of Platypleura in having the timbal cover of which inner corner is continuously rounded (not angled) to the anterior margin of the abdominal tergite 2. Description of male (Fig. 1). Head light green to greenish ochraceous with the following black marks: a median large spot enclosing ocelli; two pairs of longitudinally arranged small spots on the sides of the median spot, of which both of each pair are nearly connected to each other; a pair of oblique marks between the anterior one of the longitudinally arranged small spots and compound eyes; and a pair of small marks or fasciae on supra antennal plates. Distance between lateral ocelli and compound eyes slightly wider than twice the distance between two lateral ocelli. Postclypeus not swollen. Antenna mostly dark brown except fuscous pedicel. Ventral part of head light green to greenish ochraceous with black marks and densely covered with silvery hairs and white pollinosity. Postclypeus with transverse fasciae along transverse grooves, of which medial ends (or paramedical ends in case of anterior fasciae) are continued to each other. Anteclypeus with a pair of large marks, which occupy most part of anteclypeus except a narrow longitudinal median part. Rostrum black apically; slightly passing posterior margin of hind coxae. Lorum with a patch on posterior part. Gena with a broad transverse fascia between antenna and compound eye. Pronotum light green to greenish ochraceous. Inner area of pronotum with indistinct narrow brown patches on paramedian and
lateral fissures. Pronotal collar with a pair of transversely arranged medial tiny spots on anterior margin and dark brown patches on lateral submargin. Lateral pronotal collar broadly developed and angulated with an angle of about 100°. Mesonotum green to greenish ochraceous with the following marks: a median longitudinal black fascia, which is broadest at about center of mesonotum and sometimes elongated posteriad to touch anterior margin of cruciform elevation; a pair of small roundish black spots enclosing scutal depressions; a pair of obconical large black spots on submedian sigilla; and a pair of longitudinal long triangular castaneous patches on lateral sigilla, of which anterior ends reach anterior margin of mesonotum. Cruciform elevation ochraceous with delicate transverse fascia along anterior margin. Ventral part of thorax light green to greenish ochraceous with black marks and densely covered with silvery hairs and white pollinosity. Legs ochraceous to greenish ochraceous. Tarsi and pretarsal claws with darker parts. Fore wing venation green basally and reddish brown apically. Fore wing entirely furnished with gray and brownish cloudy marks. Hind wing ochraceous with a fuscous band along outer margin and a large fuscous spot covering over about apical halves of costal, radial, and medial cells and basal areas of apical cells 1, 2, 3, 4, and 5, and sometimes 6, which is continued with the band anteriorly. Operculum greenish ochraceous; semicircular, transversely wide; short, reaching posterior margin of sternite II. Two opercula slightly or not overlapping medially. Abdomen obconical, short, shorter than distance from head to cruciform elevation; black with greenish ochraceous caudal margin on each of tergites 2–7. Tergite 8 entirely black. Timbal cover greenish ochraceous to ochraceous; semicircular, wider than long. Timbal slightly exposed paramedially in dorsal view. Inner corner of timbal cover continuously rounded to anterior margin of tergite 2. Ventral part of abdomen dark ochraceous. Male genitalia (Fig. 2): Pygofer ellipsoidal with acuminate apex (wide dorsal beak), but rather triangular at about upper half, in ventral view. Uncus not bifurcate, thick in lateral view, with apex roundly expanded downward. Aedeagus not tapering down to apex. Upper and basal lobes of pygofer not prominent. Distribution. Philippines (Panay). Tribe Cryptotympanini Genus Cryptotympana Stål, 1861 Type species: Tettigonia atrata Fabricius, 1775. Cryptotympana Stål, 1861: 613. Cryptotympana socialis Hayashi, 1987 Cryptotympana socialis Hayashi, 1987: 13 [TL: Antique, Isl. Panay]. Distribution. Philippines (Panay, Cebu, and Negros). Tribe Cicadini Subtribe Dundubiina Genus Orientopsaltria Kato, 1944 Type species: Dundubia duarum Walker, 1857. Orientopsaltria Kato, 1944: 6. Orientopsaltria fuliginosa (Walker, 1850) Dundubia fuliginosa Walker, 1850: 60 [TL: Philippine Islands]. Orientopsaltria fuliginosa: Duffels and Zaidi, 2000: 275.
Fig. 2. Male pygofer of Platypleura lourensi sp. nov. (A) ventral view. (B) lateral view.
Material examined. 1 male, 2 females, Panay, Iloilo, Maasin, riverbed forest, replant, 100 m, 10°53.882′N 122°24.595′E, 29 VIII 2008, J.H. Lourens (IRSNB); 5 males, 9 females, Panay, Iloilo, E. of Mt. Baloy, Bgy Supanga, 32 km W. of Calinog, 570 m, 11°09.460′N 122°21.279′E, 30–31 VIII 2008, J.H. Lourens (IRSNB). Distribution. Philippines (on most Philippine islands including Panay).
Y.J. Lee / Journal of Asia-Pacific Entomology 12 (2009) 293–295
295
developed. Fore femur with primary, secondary, and two small subapical spines. Wings hyaline without infuscation. Operculum short and wide, not reaching posterior margin of sternite II. Abdomen slightly longer than distance from head to cruciform elevation. Timbal cover absent. Pygofer nearly diamond-shaped in ventral view. Upper lobe of pygofer absent. Dorsal beak dull, not acute, triangular. Median lobe of uncus triangular, slender, and very long. A pair of clasper-like processes protruding downward. Etymology. The generic name is a feminine noun derived from the Korean ggoma- and the Latin psalta, meaning “a small musician”. Ggomapsalta vernalis (Distant, 1916) (Fig. 3) Pauropsalta vernalis Distant, 1916: 204 [TL: Mt. Makiling, Luzon, Philippine Islands]. Material examined. 1 male, Panay, Iloilo, E. of Mt. Baloy, Bgy Supanga, 32 km W. of Calinog, 570 m, 11°09.460'N 122°21.279′E, 30–31 VIII 2008, J.H. Lourens (IRSNB). Distribution. Philippines (Luzon and Panay). Remarks. This species is newly recorded from Panay. Acknowledgements
Genus Dundubia Amyot and Audinet-Serville, 1843 Type species: Tettigonia vaginata Fabricius, 1787. Dundubia Amyot and Audinet-Serville, 1843: 470.
I am indebted to Dr. Jerome Constant (Institut royal des Sciences naturelles de Belgique, Brussels) for the loan of specimens. I am grateful to anonymous reviewers for suggestions that improved the manuscript. This work benefited from support from the University of Connecticut and the National Science Foundation under Grant Numbers DEB 05-29679 and DEB 07-20664. Any opinions, findings, and conclusions or recommendations expressed in this material are those of the author and do not necessarily reflect the views of the NSF.
Dundubia vaginata (Fabricius, 1787) Tettigonia vaginata Fabricius, 1787: 266 [TL: Sumatra].
References
Fig. 3. Ggomapsalta vernalis (Distant, 1916), male, Iloilo, Panay (IRSNB). (A) dorsal view. (B) ventral view. (C) ventral view of pygofer. (D) lateral view of pygofer.
Material examined. 3 males, 1 female, Panay, Iloilo, E. of Mt. Baloy, Bgy Supanga, 32 km W. of Calinog, 570 m, 11°09.460′N 122°21.279′E, 30–31 VIII 2008, J.H. Lourens (IRSNB). Distribution. S. China, Philippines (on most Philippine islands including Panay), Malay Archipelago, S. Myanmar, and India. Remarks. This species is newly recorded from Panay. Subfamily Cicadettinae Tribe Cicadettini Genus Ggomapsalta gen. nov. Type species: Pauropsalta vernalis Distant, 1916. Diagnosis. This genus is similar to Pauropsalta Goding and Froggatt, 1904 in having the following common features: a wide head, which is wider than the base of the mesonotum; hyaline wings, which have no infuscation; and the hind wing, which has only five apical cells (Figs. 3A, B). But this genus is distinguished by the nearly unicolor body and the unique structure of the male genitalia (Figs. 3C, D): upper lobe of pygofer is absent; dorsal beak is dull, not acute, triangular; median lobe of uncus is triangular, slender, and very long; a pair of clasper-like processes protrude downward. Description. Body tiny, about 12 mm long. Head including eyes wider than base of mesonotum. Distance between lateral ocelli and compound eyes slightly wider than distance between two lateral ocelli. Postclypeus scarcely swollen. Anterolateral pronotal collar not
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