Foraging CHAPTER OUTLINE 9.1 Introduction 254 9.2 Diet Choice and Food Selection 255 What Is an Adequate Diet? 255 Essential Nutrients 256 Digestion of Cellulose 256
9.3 How Animals Get Food 257 Trophic Strategies and Styles of Foraging 257 Styles of Herbivory 257 Overcoming Plant Defenses 259 Active Hunters 260 Sit and Wait Predation 262 Omnivory 262 Mimicry and Luring 262 Cooperative Hunting 263 Uninvited Guests during Foraging 264
9.5 Manipulation of Prey 265 9.6 Parasitic Life Cycles 266 9.7 Foraging and Optimality Theory 268 9.8 Optimal Patch Choice 270 The Marginal Value Theorem 270 Central Place Foraging 272 Risk Sensitivity 272
9.9 Optimal Prey Choice 274 9.10 Nutritional Constraints 275 Summary 276 Study Questions 277 Further Reading 277
LEARNING OBJECTIVES Studying this chapter should provide you with the knowledge to: l See how dietary requirements, natural selection for dietary specialization, and the evolution of foraging behavior interact. l Understand the diversity of dietary strategies employed by animals. l Be able to use optimality theory to test and refine hypotheses about important resources—the currencies of animal lives. In the study of foraging, this is frequently energy. l Understand the limits of optimality theory; animals are not expected to optimize every behavior. These constraints are the same as those that make adaptation and natural selection works in progress rather than finished portraits.