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Notes on five little known genera of Lecithoceridae (Lepidoptera), with three new species from Thailand
Journal of Asia-Pacific Entomology 12 (2009) 261–267
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Journal of Asia-Pacific Entomology j o u r n a l h o m e p a g e : w w w. e l s e v i e r. c o m / l o c a t e / j a p e
Notes on five little known genera of Lecithoceridae (Lepidoptera), with three new species from Thailand Kyu-Tek Park a,b,⁎, Chunsheng Wu c a b c
The Korean Academy of Science and Technology, Republic of Korea McGuire Center for Lepidoptera and Biodiversity, Florida Museum of Natural History, University of Florida, Gainesville, FL 32611, USA Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China
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Article history: Received 3 April 2009 Revised 3 June 2009 Accepted 9 June 2009 Keywords: Taxonomy Lecithoceridae Procharista Urolaguna Eurodachtha Tegenocharis Trichoboscis New species
a b s t r a c t Six species belonging to five little known genera of Lecithoceridae (Gelechioidea); Procharista Meyrick, Urolaguna Wu, Eurodachtha Gozmány, Tegenocharis Gozmány, and Trichoboscis Meyrick, are reported with description of morphological characters including their venation, which were not originally described in detail. Among them, three species of the two genera, Procharista ranongensis sp. nov., P. spectabilosa sp. nov., and Eurodachtha rotundina sp. nov., are described as new species from Thailand, and the female of Trichoboscis pansarista Meyrick is reported for the first time. © Korean Society of Applied Entomology, Taiwan Entomological Society and Malaysian Plant Protection Society, 2009 Published by Elsevier B.V. All rights reserved.
Introduction During recent years, a number of faunistic studies for the family Lecithoceridae (Gelechioidea) of Thailand and Vietnam have been conducted by Park (2001, 2002a,b, 2003a–c, 2004, 2005a,b, 2006a,b, 2007), describing more than 50 new species belonging to various genera of the family. In Lecithoceridae, about 50% of the known genera were described as monotypic and most of them are still known with a single species or only few species. In this study, authors found six species belonging to five little known genera of Lecithoceridae; Procharista Meyrick, Urolaguna Wu, Tegenocharis Gozmány Eurodachtha Gozmány, Eurodachtha Gozmány, and Trichoboscis Meyrick, from Thailand or Vietnam. Among them, three new species are described and the other three known species are redescribed with some morphological characters, including the venation and the genitalia, which were not originally described in detail. The genus Procharista has been known as monotypic, but two additional species are newly found inThailand and described, with more detaileddescriptionof thegenusonmorphologicalcharacters,including
⁎ Corresponding author. McGuire Center for Lepidoptera and Biodiversity, Florida Museum of Natural History, University of Florida, Gainesville, FL 32611, USA. E-mail addresses: [email protected] (K.-T. Park), [email protected] (C. Wu).
the venation and the genitalia. The genus Urolaguna is also a monotypic genus,withasingleknownspecies,U.heosaWufromChina.Thisisthefirst reportfromoutsideofChinasinceitwasdescribed.ThegenusTegenocharis wasdescribedfromNepal,basedonasinglemale,andonlyoneadditional species T. striatus Wu is known from China. The genus Eurodachtha has beenknownonlyintheMediterraneansubregion,andthedescriptionofa newspeciesfromThailandisthefirstreportfromtheOrientalRegion.The genus Trichoboscis which comprises two known species, is easily distinguished from its allies by the antenna with a notch at basal 1/5 and the unique shape of the labial palpus of the male: the 2nd segment with a long scales-tuft dorsally and the 3rd segment short, stout. Since the type species,T. pansarista MeyrickandT. crocosema(Meyrick)weredescribed from India, the latter has also been reported from China. The female of T. pansaristaisknownforthefirsttime,andthespeciesisnewlyreportedfrom ThailandorVietnamsinceitwasdescribed. Materials and methods Material examined includes specimens deposited in the Osaka Prefecture University (OPU), which were collected by Japanese microlepidopterists during their expeditions in 1981, 1985 and 1987. For the descriptions of new species, colors follow Kornerup and Wanscher (1983). Types will be deposited in the OPU, Japan, on an indefinite loan from Thailand.
1226-8615/$ – see front matter © Korean Society of Applied Entomology, Taiwan Entomological Society and Malaysian Plant Protection Society, 2009 Published by Elsevier B.V. All rights reserved. doi:10.1016/j.aspen.2009.06.002
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Systematics Genus Procharista Meyrick, 1922 Procharista Meyrick, 1922. Zool. Med. Lepid. 7: 82; Meyrick, 1925: 225. Type species: P. sardonias Meyrick, 1922: 225. TL: Java. The genus is known as a monotypic genus. The genus is characterized by the wing shape, especially the trapezoidal-ovate hind wing with well developed hair-pencils on costa basally; the venation with M1 and M2 coincident, CuA1 and CuA2 coincident in the forewing, and M2 absent in the hindwing; and the male genitalia with gnathos almost atrophied or very small and valva with outward directed strong process on ventral margin. The venation is redescribed in the following new species. The abdominal tergites have spinous zones. The distributional range includes Indonesia (Java) and Thailand (new record). Procharista ranongensis sp. nov. (Figs. 1, 1a, 1b, 12, 12a–c) Types. Holotype: ♂, Thailand, Ranong, Na Kha, 250 m, 29–31 X 1985 (Moriuti, Saito, Arita), gen. slide no. 5827/Park. Diagnosis. The new species is distinguished from the type species, P. sardonias Meyrick, by the smaller size, 2nd segment of labial palpus without dark scales at apex, and only part of veins marked by cloudy whitish lines, without dark blue line below costa on basal 1/5, and uniform clothed with brownish short hair-like scales on the hindwing. Descritpion. Wingspan, 13.5 mm. Head covered with metallic, coppery-purple scales on vertex medially; sidetufts with smoothly appressed pale orange to pale grayish orange scales. Scape of antenna
elongated, pale grayish orange dorsally, orange white ventrally; flagellum pale grayish orange, without distinct annulations. Labial palpus slender, strongly upturned; 2nd segment of labial palpus slender, shiny orange white on both surfaces; 3rd segment as long as 2nd, dark brown ventrally. Tegula and thorax yellowish brown. Forewing elongate, more or less lanceolate, with pointed apex; ground color uniform yellowish brown, clothed with cloudy whitish scales along veins, especially R and M-veins; venation nearly same as that of the following new species; fringe yellowish brown on termen, dark brown near tornus. Hindwing broad, nearly trapezoidal-ovate, uniform clothed with brownish short hair-like scales, with well developed orange white hair-pencils along costa basally; apex rounded; termen slightly oblique; fringe concolorous; venation with Rs and M1 nearly connate; M2 absent; M3 and CuA1 short stalked; cell partly closed. Hind tibia with brownish yellow hair-like scales dorsally, with dark brown scales preapically; inner spur dark brown; tarsi speckled with dark brown scales. The abdominal tergites weakly sclerotized, sparsely spinous; the 8th caudal margin emarginated at middle, with medial lobes. Female unknown. Male genitalia (Figs. 12, 12a, c). Uncus-lobes short, semiovate. Gnathos short, small, weakly sclerotized (Fig. 12c). Tegumen short, about 1/2 length of vinculum. Costal bar heavily sclerotized, slightly angled at middle. Valva with basal half trapezoidal, broader toward distally; cucullus slender, elongate, sword-shaped, longer than basal part, with heavily sclerotized, short, spine-like process at near base on ventral margin, with dense setae on surface; sacculus bifurcated basally. Juxta concave, weakly sclerotized on caudal margin, with small median plate anteriorly. Aedeagus stout, as long as valva; cornutus horn-shaped, heavily sclerotized, about 2/5 length of aedeagus.
Figs. 1–3. Adults (a: dorsal view of head; b: labial palpus). 1, Procharista ranongensis sp. nov., holotype; 2, Procharista spectabilosa sp. nov., holotype; 3, Urolaguna heosa Wu.
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Distribution. Thailand (South). Etymology. The species name is derived from the type locality. Procharista spectabilosa sp. nov. (Figs. 2, 2a, 2b, 8, 13, 13a–c) Types. Holotype: ♂, Thailand, Ranong, Na Kha, 250 m, 29–31 X 1985 (Moriuti, Saito, Arita), gen. slide no. 5855/Park. Paratype. 1♂, same data as the holotype, wing prep. no. 5845/Park. Diagnosis. This new species is very similar to the preceding new species, but can be distinguished as follows: vertex more metallic purplish; forewing veins nearly not marked by cloudy whitish lines, apex less acute; hindwing more trapezoidal, with obtuse apex, M3 + CuA1 with longer stalk. The male genitalia are also similar, but the gnathos is nearly atrophied, the ventral process of valva is more slender and ventro-inwardly directed, and the caudal margin of juxta is concaved into V-shape. Description. Wingspan, 12.5–13.0 mm. Head covered with metallic, coppery-purple scales dorsally, more broadly covered than that of P. ranongensis; sidetufts weakly presented with pale orange scales. Antenna and labial palpus nearly same as those of the preceding new species. Tegula and thorax yellowish brown. Forewing elongate, more or less lanceolated; apex less pointed; ground color uniform yellowish brown; veins not cloudy whitish along veins on surface; forewing venation (Fig. 8) with R2 arising from 3/5 length of R1 – R3; R3 stalked with R4 + 5 near basal 1/4; R4 and R5 stalked near middle; R5 to apex; M1 far from R3 + 4 + 5 at base; M2 and M3 coincident; CuA1 and CuA2
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coincident; cell very narrowed at basal half; weakly closed; fringe yellowish brown, concolorous. Hindwing nearly trapezoidal-ovate, uniform clothed with brownish short hair-like scales on surface, with well developed orange white hair-pencils along costa basally; apex rounded; termen slightly oblique; fringe concolorous; venation with Rs and M1 nearly connate; M2 absent; M3 and CuA1 stalked at basal 1/4; cell partly closed. Hind tibia also similar to that of the preceding species. The abdominal tergites weakly sclerotized, sparsely spinous; 8th caudal margin emarginated at middle, medial lobes less acute (Fig. 13b). Male genitalia (Figs. 13, 13a, c). Uncus-lobes not divided, crescent. Gnathos minute, almost atrophied (Fig. 13c).Tegumen as long as vinculum. Costal bar heavily sclerotized, without median angle. Basal half of valva produced at lower angle with round apex; cucullus narrower, longer than that of preceding species, longer than basal part with nearly parallel sides, dense setosed on surface; ventral process heavily sclerotized, as long as width of cucullus, directed ventroinwardly. Juxta with V-shaped caudal margin, with small median plate anteriorly. Aedeagus proportionally slender, small, about 1/2 length of valva; cornutus horn-shaped, about 2/5 length of aedeagus. Distribution. Thailand (South). Etymology. The name is derived from Latin, spectabil ( = remarkable) and Latin suffix, osus(a). Genus Urolaguna Wu, 1994 Sinozoologia 11: 132–133. Type species: Urolaguna heosa Wu, 1994. TL: Jianxi, China.
Figs. 4–7. Adults (a: dorsal view of head; b: labial palpus). 4, Tegenocharis tenebrans Gozmány; 5, Tegenocharis striatus Wu, holotype; 6, Eurodachtha rotundina sp. nov., holotype; 7, Trichoboscis pansarista Meyrick.
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Figs. 8–11. Wings venation. 8, Procharista spectabilosa sp. nov.; 9, Urolaguna heosa Wu; 10, Tegenocharis tenebrans Gozmány; 11, Eurodachtha rotundina sp. nov.
Urolaguna Wu is a monotypic genus which was established, as separated from Eurodachtha Gozmány. According to the original description, it was differentiated from Eurodachtha by the venation: PCu well presented, M2 free from M3, nearly parallel, CuA1 and CuA2 stalked in the forewing. The description of the venation (Fig. 9) is given in the following species. The male genitalia also have some different shape of the valva and aedeagus from Eurodachtha. The abdominal tergites have spinous zones dorsally. The distributional range includes China and Thailand (new record).
The wing venation of the genus is similar to that of Lecithocera, but it is distinguished by having the spinous zones on the abdomen dorsally. It is more or less related to Hoenea Gozmány and Siderostigma Gozmány, but also differs from them by the elongate forewing, with R3 stalked R4 + 5. The forewing is slender, narrow and elongated, with M1 far from the stalk of R3 + 4 + 5 and CuA1 and CuA2 far from each other. The abdominal tergites have spinous zones dorsally, with removable spines. The distributional range of the genus includes Nepal, Thailand (new record), and China, with two known species.
Urolaguna heosa Wu, 1994 (Figs. 3, 3a, 3b, 9, 14, 14a, 14b, 15) Urolaguna heosa Wu, 1994, Sinozoologia 11: 133; 1987: 196. TL: Jiangxi. Type in the Institute of Zoology (IZCAS), Beijing.
Tegenocharis tenebrans Gozmány (Figs. 4, 4a, 4b, 10, 16, 16a, b) Tegenocharis tenebrans Gozmány, 1973, Khumbu Himal. 4: 430; Gozmány, 1978: 136. TL: Nepal. Type in the HNHM, Budapest.
Diagnosis. Wingspan, 13.5–16.5 mm. The forewing shape and ground color are similar to the preceding species of the genus Procharista, but narrower and lanceolate with acute apex. The hindwing has more oblique termen and the costa anteriorly expanded at basal half. Description of venation. Venation with R2 closer to R1 at base; R3 free from R4 + 5; R4 and R5 stalked near middle; R5 terminated to termen; M2 free from M3 at base, nearly parallel; CuA1 and CuA2 stalked at basal 1/5; cell closed; apex acute; termen extremely oblique. Hindwing venation with Rs and M1 short stalked; M2 absent; M3 and CuA1 connate. Male genitalia (Figs. 14a, b). Described and illustrated by Wu (1994, Fig. 25; 1997, Pl. 12, Fig. 1). Female genitalia (Fig. 15). Described and illustrated by Wu (1994, Fig. 33; 1997, Pl. 25, Fig. 2). Material examined. 2♂, 1♀, Thailand, Chiang Mai, Fang, 450 m, 29– 31 X 1985 (Moriuti, Saito, Arita), gen slide no. 5839/Park (♂), 5849/ Park (♀); 1♂, Nakhon Nayok, Khao Yai, 800 m, 19 VI 1983 (Kuroko, Moriuti, Arita, Yoshimoto ), wing prep. no. 5841/Park; 1♂, Nakhon Nayok, Khao Yai, 800 m, 21 IX 1987 (Moriuti, Saito, Arita, Yoshiyashu). Distribution. China (Jiangxi), Thailand (new record; Chiang Mai, Nakhon Nayok).
Diagnosis. Wingspan,12.5–13.0 mm. Forewing ground color is grayish orange to pale brownish orange. The coloring is much paler than the type specimen, and also some little differences of the forewing venation are found between the type and specimens of Thailand, especially the distance of R2 from R3 at base, length of stalk of R3 and R4 + 5. However, it is considered that these differences can be a geographical variation. The Chinese species, T. striatus differs from the type species by the clearly presented discal spots and whitish lines along veins on the forewing (Fig. 5). The male genitalia are also very similar, but gnathos is narrower and longer, and the distal part of valva is proportionally shorter. Female unknown. Male genitalia (Figs. 16, 16a, b). Described and illustrated by Gozmány (1978, Taf. 36, Fig. 80). Material examined. 4♂, Chiang Mai, Doi Pui, 1300 m, 1–4 IX 1987 (Moriuti, Saito, Arita, Yoshimoto), gen. slide no. 5828/Park. Distribution. Nepal, Thailand (new record; Chiang Mai).
Genus Tegenocharis Gozmány, 1973 Tegenocharis Gozmány, 1973, Khumbu Himal. 4: 429; 1978: 135: Wu, 1997: 199. Type species: Tegenocharis tenebrans Gozmány, 1973.
Genus Eurodachtha Gozmány, 1978 Microlep. Palaearctica 5: 151. Type species: Lecithocera pallicornella Staudinger, 1859. TL: Spain. Eurodachtha Gozmány has narrow, elongated forewing, with all veins: R3 free from R4 + 5, M2 and M3 free; CuA1 and CuA2 far from each other; hindwing with M2 atrophied, often with only vestige (originally described as M2 and M3 coincident); M3 and CuA1 short
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Figs. 12–15. Male genitalia, abdominal segments, and female genitalia. 12, Procharista ranongensis sp. nov. sl. No. 5827; 12a, ditto, aedeagus; 12b, ditto, abdominal segment; 13, Procharista spectabilosa sp. nov., sl. No. 5855; 13a, ditto, aedeagus; 13b, ditto, abdominal segment; 13c, ditto, enlargement of gnathos; 14, Urolaguna heosa Wu, sl. No. 5839; 14a, ditto, aedeagus; 14b, ditto, abdominal segment; 15. Female genitalia of Urolaguna heosa Wu, sl. No. 5849.
stalked or connate; CuA2 arising from middle or beyond (Fig. 11). Abdominal tergites have sparsely spiniform setae which are easily removable. Five species were known only in the Mediterranean subregion, but no species has been known from the Oriental Region. The following new species is reported for the first time from the Oriental Region. Eurodachtha rotundina sp. nov. (Figs. 6, 6a, 6b, 11, 17, 17a, b) Type. Holotype. ♂, Thailand, Nakhon Noyak, Khao Yai, 800 m, 20 VI 1983 (Kuroko, Moriuti, Arita, Yoshiyasu), gen. slide no. 5829/Park, wings prep. no. 5851/Park.
Diagnosis. The species is more or less similar to E. nigralba Gozmány, which was described from Syria, but differs from the latter by the yellowish brown forewing and the different shape of the valva of the male genitalia, with strongly bent cucullus. Description. Wingspan, 12.5 mm. Head yellowish brown, with grayish orange erect scales laterally. Scape of antenna elongated, slightly broadened toward apex, grayish orange dorsally, orange white ventrally; flagellum yellowish brown, with indistinct dark brown annulations; apex pale orange gray. Labial palpus slender, strongly upturned; second segment slender, shiny orange white on both surfaces; 3rd segment as long as 2nd, dark brown ventrally. Tegula and thorax yellowish brown. Forewing elongate, lanceolate, with acute
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Figs. 16–19. Male genitalia, abdominal segments, and female genitalia. 16, Tegenocharis tenebrans Gozmány, sl. No. 5828; 16a, ditto, aedeagus; 16b, ditto, abdominal segment; 17, Eurodachtha rotundina sp. nov., sl. No. 5829; 17a, ditto, aedeagus; 17b, ditto, abdominal segment; 18, Trichoboscis pansarista Meyrick, sl. No. 5840; 18a, ditto, aedeagus; 18b, enlargement of aedeagus, sl. No. 5087; 19, Female genitalia of Trichoboscis pansarista Meyrick, sl. No. 5857.
apex; ground color uniform metallic yellowish brown, without any markings; costa nearly straight prior to 4/5 length; termen strongly oblique; fringe concolorous; venation with R1 arising before middle of cell; R2 arising nearly from middle; R3 free from R4 + 5; R4 and R5 stalked near middle; R5 terminated to apex; M1 far from R4 + 5 at base; M2 nearly parallel to M1; M2 and M3 free; CuA1 and CuA2 nearly connate, proportionally short; cell closed; apex acute; termen extremely oblique. Hindwing pale orange gray; costa nearly straight; apex acute; termen oblique, slightly concave medially; venation with
Rs and M1 stalked at near middle; M2 atrophied, with only vestige; M3 and CuA1 connate; CuA2 arising from middle of cell. Hind tibia with pale grayish orange hair-like scales dorsally; spurs proportionally long, longer one about 1/2 length of hind tibia; tarsi concolorous. Abdominal tergites have sparsely spiniform setae dorsally. Female unknown. Male genitalia (Figs. 17, 17a, b). Uncus-lobes short, semiovate. Gnathos short, prorptionally small. Costal bar sharply angled at middle. Valva broad at basal 1/3, with large cavity on surface; distal 2/ 5 nearly parallel, curved distally, dense setosed, with rounded apex;
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costa strongly concaved, with dense setae on surface; sacculus broad, with long setae. Juxta shield-shaped; caudal margin concave with narrowly produced laterocaudal processes. Aedeagus stout, as long as valva, with various length of short strong spines beyond middle. Distribution. Thailand (Nakhon Nayok). Genus Trichoboscis Meyrick, 1929 Exot. Microlep. 3: 526; Gozmány, 1978: 176. Wu, 1997: 198. Type species: Trichoboscis pansarista Meyrick, 1929. Trichoboscis Meyrick is easily distinguished from its allies by the antenna with a notch at basal 1/5 and the unique shape of the labial palpus of the male: the 2nd segment with a long scales-tuft dorsally and the 3rd segment short, stout. The labial palpus of the female is normally recurved. Abdominal tergites have spinous zones dorsally. Two species are known from India and China. Trichoboscis pansarista Meyrick, 1929 (Figs. 7, 7a, 7b, 18, 18a, b, 19) Trichoboscis pansarista Meyrick, 1929, Exot. Microlep. 3: 526; Clarke, 1965: 255; Gozmány, 1978: 176. TL: Sikkim, India. Type in the Natural History Museum (BMNH), London. Diagnosis. Forewing length 15 mm. The male genitalia of the species are very similar those of T. (Clarke, 1965, Pl. 61, Figs. 2–2b), but can be distinguished by the less slender valva and more thorn-like spines in the aedeagus. The female genitalia also differs by a single plate of signum in T. pansarista Meyrick, instead of three plate of signa in T. crocosema Meyrick. Male genitalia (Figs. 18, 18a, b). Illustrated by Clarke (1965, Pl. 127, Figs. 1c–d), Gozmány (1978, Taf. 43, Fig. 106). Female genitalia (Fig. 19). Eighth abdominal segment weakly sclerotized, slightly emarginated at middle on caudal margin. Apophyses anteriores about 1/2 length of apophyses posteriors. Antrum memberanous. Ductus bursae short, funnel-like, about 2/3 length of corpus bursae. Corpus bursae ovate, bearing numerous, very short corn-shaped spines with broad basal part on internal surface; signum small, semiovate, and radiated. Material examined. 1♂, Vietnam, Cuc Phuong Nat. Park, 300 m, 2 V 2005 (KT Park and SR Kim), gen. slide no. CIS-5087/Park; 1♂, same locality as above, 30 IV–1V 2005 (KT Park and SR Kim); 7♂, Vietnam, Bac Khan Prov., Ba Be National Park, 240 m, 26–28 VII 2006 (Park, Chae, and Cuong); 1♂, same locality as above, 21–26 Jun 2008 (JB Heppner); 2♂, Thailand, Nakhon Nayok, Khao Yai, 800 m, 21–22 VI 1983 (Kuroko, Moriuti, Saito, Arita, Yoshiyasu); 1♂, 1♀, Thailand, Chiang Mai, Khao Soi Dao, 480 m, 24–25 VIII 1987 (Moriuti, Saito, Arita, Yoshimoto), gen. slide no. 5840/Park(♂) and 5857/Park(♀); 1♂, same locality, 22 IX 1987 (Moriuti, Saito, Arita, Yoshiyasu).
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Distribution. India (North), Thailand (new record; North and Central), Vietnam (new record; North). Acknowledgments Authors are grateful to T. Hirowatari, Osaka Prefecture University, Osaka, Japan, for the loan of valuable specimens for this study, and M Thomas, Florida State Collection of Arthropods, Division of Plant Industry, FDACS, Gainesville, FL, USA, for his allowance in using all available equipments, including Automontage microscope. A part of the financial support was given by the National Institute of Biological Resource, Korea. References Clarke, J.F.G., 1965. Catalogue of the type specimens of Microlepidoptera in the British Museum (Natural History) described by Edward Meyrick. Vol. 5, Vol. 5. London, 581 pp. Gozmány, L., 1973. Symmocid and Lecithocerid moths (Lepidoptera) from Nepal. Khumbu Himalaya 4 (3), 413–444. Gozmány, L., 1978. Lecithoceridae. In: Amsel, H.G., Gregor, F., Reisser, H. (Eds.), Microlepidoptera Palaeartica, Vol. 5. Georg Fromme and Co., Wien. 306 pp. Kornerup, A., Wanscher, J.H., 1983. Methuen Handbook of Colour. 3rd ed. 252 pp. Methuen and Co. Ltd., London. Meyrick, E., 1922. New Micro-Lepidoptera. Zool. Meded. 7, 80–89. Meyrick, E. 1929. Exotic Microlepidoptera 3: 521–526. Marloborough, Wilts. Park, K.T., 2001. A new species of the genus Tisis (Lepidoptera: Lecithoceridae) from Thailand. Korean J. Syst. Zool. 17 (1), 59–63. Park, K.T., 2002a. Taxonomic review of the genus Torodora Meyrick in Thailand, with descriptions of fifteen new species (Lepidoptera: Lecithoceridae). Insecta Koreana 19 (2), 147–166. Park, K.T., 2002b. Four new species of the genus Antiochtha Meyrick (Lepidoptera: Lecithoceridae) from Thailand. J. Asia-Pacific Entomol. 5 (2), 139–143. Park, K.T., 2003a. Two new species of the genus Tisis from Thailand and notes on Tisis elegans (Insecta: Lepidoptera: Lecithoceridae). Species Divers. 8, 267–274. Park, K.T., 2003b. Three new species of the genus Hygroplasta Meyrick (Lepidoptera, Lecithoceridae) from the Philippines and Thailand. J. Asia-Pacific Entomol. 6 (2), 131–135. Park, K.T., 2003c. Thubana species (Lepidoptera, Lecithoceridae) in Thailand, with descriptions of twelve new species. J. Asia-Pacific Entomol. 6 (2), 137–150. Park, K.T., 2004. Genus Homaloxestis Meyrick of Thailand, with description of five new species (Lepidoptera, Lecithoceridae). J. Asia-Pacific Entomol. 7 (1), 33–43. Park, K.T., 2005a. Four new species of the genus Frisilia Walker from Thailand (Lepidoptera: Lecithoceridae, Lecithocerinae). Tinea 18 (Suppl. 3), 55–62. Park, K.T., 2005b. Genus Lecithocera Herrich-Shäffer of Thailand (I): descriptions of three new species (Lepidoptera, Lecithoceridae). J Asia-Pacific Entomo. 8 (3), 233–237. Park, K.T., 2006a. Genus Lecithocera of Thailand (II): descriptions of four new species (Lepidoptera, Lecithoceridae). Tinea 19 (2), 98–103. Park, K.T., 2006b. Genus Lecithocera of Thailand (III): new records of five species of the genus (Lepidoptera, Lecithoceridae). J. Asia-Pacific Entomol. 9 (4), 313–316. Park, K.T., 2007. Three new species of Torodora Meyrick from Thailand (Lepidoptera, Lecithoceridae). Shilap Revista Lepidop. 35, 23–28. Wu, C., 1994. The Lecithoceridae (Lepidoptera) of China, with description of new taxa. Sinzoologica 11, 123–154. Wu, C., 1997. Lepidoptera, Lecithoceridae. Fauna Sinica, Insecta., Vol. 7. Science Press, Beijing, pp. 1–302.
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Journal of Asia-Pacific Entomology j o u r n a l h o m e p a g e : w w w. e l s e v i e r. c o m / l o c a t e / j a p e
Notes on five little known genera of Lecithoceridae (Lepidoptera), with three new species from Thailand Kyu-Tek Park a,b,⁎, Chunsheng Wu c a b c
The Korean Academy of Science and Technology, Republic of Korea McGuire Center for Lepidoptera and Biodiversity, Florida Museum of Natural History, University of Florida, Gainesville, FL 32611, USA Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China
a r t i c l e
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Article history: Received 3 April 2009 Revised 3 June 2009 Accepted 9 June 2009 Keywords: Taxonomy Lecithoceridae Procharista Urolaguna Eurodachtha Tegenocharis Trichoboscis New species
a b s t r a c t Six species belonging to five little known genera of Lecithoceridae (Gelechioidea); Procharista Meyrick, Urolaguna Wu, Eurodachtha Gozmány, Tegenocharis Gozmány, and Trichoboscis Meyrick, are reported with description of morphological characters including their venation, which were not originally described in detail. Among them, three species of the two genera, Procharista ranongensis sp. nov., P. spectabilosa sp. nov., and Eurodachtha rotundina sp. nov., are described as new species from Thailand, and the female of Trichoboscis pansarista Meyrick is reported for the first time. © Korean Society of Applied Entomology, Taiwan Entomological Society and Malaysian Plant Protection Society, 2009 Published by Elsevier B.V. All rights reserved.
Introduction During recent years, a number of faunistic studies for the family Lecithoceridae (Gelechioidea) of Thailand and Vietnam have been conducted by Park (2001, 2002a,b, 2003a–c, 2004, 2005a,b, 2006a,b, 2007), describing more than 50 new species belonging to various genera of the family. In Lecithoceridae, about 50% of the known genera were described as monotypic and most of them are still known with a single species or only few species. In this study, authors found six species belonging to five little known genera of Lecithoceridae; Procharista Meyrick, Urolaguna Wu, Tegenocharis Gozmány Eurodachtha Gozmány, Eurodachtha Gozmány, and Trichoboscis Meyrick, from Thailand or Vietnam. Among them, three new species are described and the other three known species are redescribed with some morphological characters, including the venation and the genitalia, which were not originally described in detail. The genus Procharista has been known as monotypic, but two additional species are newly found inThailand and described, with more detaileddescriptionof thegenusonmorphologicalcharacters,including
⁎ Corresponding author. McGuire Center for Lepidoptera and Biodiversity, Florida Museum of Natural History, University of Florida, Gainesville, FL 32611, USA. E-mail addresses: [email protected] (K.-T. Park), [email protected] (C. Wu).
the venation and the genitalia. The genus Urolaguna is also a monotypic genus,withasingleknownspecies,U.heosaWufromChina.Thisisthefirst reportfromoutsideofChinasinceitwasdescribed.ThegenusTegenocharis wasdescribedfromNepal,basedonasinglemale,andonlyoneadditional species T. striatus Wu is known from China. The genus Eurodachtha has beenknownonlyintheMediterraneansubregion,andthedescriptionofa newspeciesfromThailandisthefirstreportfromtheOrientalRegion.The genus Trichoboscis which comprises two known species, is easily distinguished from its allies by the antenna with a notch at basal 1/5 and the unique shape of the labial palpus of the male: the 2nd segment with a long scales-tuft dorsally and the 3rd segment short, stout. Since the type species,T. pansarista MeyrickandT. crocosema(Meyrick)weredescribed from India, the latter has also been reported from China. The female of T. pansaristaisknownforthefirsttime,andthespeciesisnewlyreportedfrom ThailandorVietnamsinceitwasdescribed. Materials and methods Material examined includes specimens deposited in the Osaka Prefecture University (OPU), which were collected by Japanese microlepidopterists during their expeditions in 1981, 1985 and 1987. For the descriptions of new species, colors follow Kornerup and Wanscher (1983). Types will be deposited in the OPU, Japan, on an indefinite loan from Thailand.
1226-8615/$ – see front matter © Korean Society of Applied Entomology, Taiwan Entomological Society and Malaysian Plant Protection Society, 2009 Published by Elsevier B.V. All rights reserved. doi:10.1016/j.aspen.2009.06.002
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Systematics Genus Procharista Meyrick, 1922 Procharista Meyrick, 1922. Zool. Med. Lepid. 7: 82; Meyrick, 1925: 225. Type species: P. sardonias Meyrick, 1922: 225. TL: Java. The genus is known as a monotypic genus. The genus is characterized by the wing shape, especially the trapezoidal-ovate hind wing with well developed hair-pencils on costa basally; the venation with M1 and M2 coincident, CuA1 and CuA2 coincident in the forewing, and M2 absent in the hindwing; and the male genitalia with gnathos almost atrophied or very small and valva with outward directed strong process on ventral margin. The venation is redescribed in the following new species. The abdominal tergites have spinous zones. The distributional range includes Indonesia (Java) and Thailand (new record). Procharista ranongensis sp. nov. (Figs. 1, 1a, 1b, 12, 12a–c) Types. Holotype: ♂, Thailand, Ranong, Na Kha, 250 m, 29–31 X 1985 (Moriuti, Saito, Arita), gen. slide no. 5827/Park. Diagnosis. The new species is distinguished from the type species, P. sardonias Meyrick, by the smaller size, 2nd segment of labial palpus without dark scales at apex, and only part of veins marked by cloudy whitish lines, without dark blue line below costa on basal 1/5, and uniform clothed with brownish short hair-like scales on the hindwing. Descritpion. Wingspan, 13.5 mm. Head covered with metallic, coppery-purple scales on vertex medially; sidetufts with smoothly appressed pale orange to pale grayish orange scales. Scape of antenna
elongated, pale grayish orange dorsally, orange white ventrally; flagellum pale grayish orange, without distinct annulations. Labial palpus slender, strongly upturned; 2nd segment of labial palpus slender, shiny orange white on both surfaces; 3rd segment as long as 2nd, dark brown ventrally. Tegula and thorax yellowish brown. Forewing elongate, more or less lanceolate, with pointed apex; ground color uniform yellowish brown, clothed with cloudy whitish scales along veins, especially R and M-veins; venation nearly same as that of the following new species; fringe yellowish brown on termen, dark brown near tornus. Hindwing broad, nearly trapezoidal-ovate, uniform clothed with brownish short hair-like scales, with well developed orange white hair-pencils along costa basally; apex rounded; termen slightly oblique; fringe concolorous; venation with Rs and M1 nearly connate; M2 absent; M3 and CuA1 short stalked; cell partly closed. Hind tibia with brownish yellow hair-like scales dorsally, with dark brown scales preapically; inner spur dark brown; tarsi speckled with dark brown scales. The abdominal tergites weakly sclerotized, sparsely spinous; the 8th caudal margin emarginated at middle, with medial lobes. Female unknown. Male genitalia (Figs. 12, 12a, c). Uncus-lobes short, semiovate. Gnathos short, small, weakly sclerotized (Fig. 12c). Tegumen short, about 1/2 length of vinculum. Costal bar heavily sclerotized, slightly angled at middle. Valva with basal half trapezoidal, broader toward distally; cucullus slender, elongate, sword-shaped, longer than basal part, with heavily sclerotized, short, spine-like process at near base on ventral margin, with dense setae on surface; sacculus bifurcated basally. Juxta concave, weakly sclerotized on caudal margin, with small median plate anteriorly. Aedeagus stout, as long as valva; cornutus horn-shaped, heavily sclerotized, about 2/5 length of aedeagus.
Figs. 1–3. Adults (a: dorsal view of head; b: labial palpus). 1, Procharista ranongensis sp. nov., holotype; 2, Procharista spectabilosa sp. nov., holotype; 3, Urolaguna heosa Wu.
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Distribution. Thailand (South). Etymology. The species name is derived from the type locality. Procharista spectabilosa sp. nov. (Figs. 2, 2a, 2b, 8, 13, 13a–c) Types. Holotype: ♂, Thailand, Ranong, Na Kha, 250 m, 29–31 X 1985 (Moriuti, Saito, Arita), gen. slide no. 5855/Park. Paratype. 1♂, same data as the holotype, wing prep. no. 5845/Park. Diagnosis. This new species is very similar to the preceding new species, but can be distinguished as follows: vertex more metallic purplish; forewing veins nearly not marked by cloudy whitish lines, apex less acute; hindwing more trapezoidal, with obtuse apex, M3 + CuA1 with longer stalk. The male genitalia are also similar, but the gnathos is nearly atrophied, the ventral process of valva is more slender and ventro-inwardly directed, and the caudal margin of juxta is concaved into V-shape. Description. Wingspan, 12.5–13.0 mm. Head covered with metallic, coppery-purple scales dorsally, more broadly covered than that of P. ranongensis; sidetufts weakly presented with pale orange scales. Antenna and labial palpus nearly same as those of the preceding new species. Tegula and thorax yellowish brown. Forewing elongate, more or less lanceolated; apex less pointed; ground color uniform yellowish brown; veins not cloudy whitish along veins on surface; forewing venation (Fig. 8) with R2 arising from 3/5 length of R1 – R3; R3 stalked with R4 + 5 near basal 1/4; R4 and R5 stalked near middle; R5 to apex; M1 far from R3 + 4 + 5 at base; M2 and M3 coincident; CuA1 and CuA2
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coincident; cell very narrowed at basal half; weakly closed; fringe yellowish brown, concolorous. Hindwing nearly trapezoidal-ovate, uniform clothed with brownish short hair-like scales on surface, with well developed orange white hair-pencils along costa basally; apex rounded; termen slightly oblique; fringe concolorous; venation with Rs and M1 nearly connate; M2 absent; M3 and CuA1 stalked at basal 1/4; cell partly closed. Hind tibia also similar to that of the preceding species. The abdominal tergites weakly sclerotized, sparsely spinous; 8th caudal margin emarginated at middle, medial lobes less acute (Fig. 13b). Male genitalia (Figs. 13, 13a, c). Uncus-lobes not divided, crescent. Gnathos minute, almost atrophied (Fig. 13c).Tegumen as long as vinculum. Costal bar heavily sclerotized, without median angle. Basal half of valva produced at lower angle with round apex; cucullus narrower, longer than that of preceding species, longer than basal part with nearly parallel sides, dense setosed on surface; ventral process heavily sclerotized, as long as width of cucullus, directed ventroinwardly. Juxta with V-shaped caudal margin, with small median plate anteriorly. Aedeagus proportionally slender, small, about 1/2 length of valva; cornutus horn-shaped, about 2/5 length of aedeagus. Distribution. Thailand (South). Etymology. The name is derived from Latin, spectabil ( = remarkable) and Latin suffix, osus(a). Genus Urolaguna Wu, 1994 Sinozoologia 11: 132–133. Type species: Urolaguna heosa Wu, 1994. TL: Jianxi, China.
Figs. 4–7. Adults (a: dorsal view of head; b: labial palpus). 4, Tegenocharis tenebrans Gozmány; 5, Tegenocharis striatus Wu, holotype; 6, Eurodachtha rotundina sp. nov., holotype; 7, Trichoboscis pansarista Meyrick.
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Figs. 8–11. Wings venation. 8, Procharista spectabilosa sp. nov.; 9, Urolaguna heosa Wu; 10, Tegenocharis tenebrans Gozmány; 11, Eurodachtha rotundina sp. nov.
Urolaguna Wu is a monotypic genus which was established, as separated from Eurodachtha Gozmány. According to the original description, it was differentiated from Eurodachtha by the venation: PCu well presented, M2 free from M3, nearly parallel, CuA1 and CuA2 stalked in the forewing. The description of the venation (Fig. 9) is given in the following species. The male genitalia also have some different shape of the valva and aedeagus from Eurodachtha. The abdominal tergites have spinous zones dorsally. The distributional range includes China and Thailand (new record).
The wing venation of the genus is similar to that of Lecithocera, but it is distinguished by having the spinous zones on the abdomen dorsally. It is more or less related to Hoenea Gozmány and Siderostigma Gozmány, but also differs from them by the elongate forewing, with R3 stalked R4 + 5. The forewing is slender, narrow and elongated, with M1 far from the stalk of R3 + 4 + 5 and CuA1 and CuA2 far from each other. The abdominal tergites have spinous zones dorsally, with removable spines. The distributional range of the genus includes Nepal, Thailand (new record), and China, with two known species.
Urolaguna heosa Wu, 1994 (Figs. 3, 3a, 3b, 9, 14, 14a, 14b, 15) Urolaguna heosa Wu, 1994, Sinozoologia 11: 133; 1987: 196. TL: Jiangxi. Type in the Institute of Zoology (IZCAS), Beijing.
Tegenocharis tenebrans Gozmány (Figs. 4, 4a, 4b, 10, 16, 16a, b) Tegenocharis tenebrans Gozmány, 1973, Khumbu Himal. 4: 430; Gozmány, 1978: 136. TL: Nepal. Type in the HNHM, Budapest.
Diagnosis. Wingspan, 13.5–16.5 mm. The forewing shape and ground color are similar to the preceding species of the genus Procharista, but narrower and lanceolate with acute apex. The hindwing has more oblique termen and the costa anteriorly expanded at basal half. Description of venation. Venation with R2 closer to R1 at base; R3 free from R4 + 5; R4 and R5 stalked near middle; R5 terminated to termen; M2 free from M3 at base, nearly parallel; CuA1 and CuA2 stalked at basal 1/5; cell closed; apex acute; termen extremely oblique. Hindwing venation with Rs and M1 short stalked; M2 absent; M3 and CuA1 connate. Male genitalia (Figs. 14a, b). Described and illustrated by Wu (1994, Fig. 25; 1997, Pl. 12, Fig. 1). Female genitalia (Fig. 15). Described and illustrated by Wu (1994, Fig. 33; 1997, Pl. 25, Fig. 2). Material examined. 2♂, 1♀, Thailand, Chiang Mai, Fang, 450 m, 29– 31 X 1985 (Moriuti, Saito, Arita), gen slide no. 5839/Park (♂), 5849/ Park (♀); 1♂, Nakhon Nayok, Khao Yai, 800 m, 19 VI 1983 (Kuroko, Moriuti, Arita, Yoshimoto ), wing prep. no. 5841/Park; 1♂, Nakhon Nayok, Khao Yai, 800 m, 21 IX 1987 (Moriuti, Saito, Arita, Yoshiyashu). Distribution. China (Jiangxi), Thailand (new record; Chiang Mai, Nakhon Nayok).
Diagnosis. Wingspan,12.5–13.0 mm. Forewing ground color is grayish orange to pale brownish orange. The coloring is much paler than the type specimen, and also some little differences of the forewing venation are found between the type and specimens of Thailand, especially the distance of R2 from R3 at base, length of stalk of R3 and R4 + 5. However, it is considered that these differences can be a geographical variation. The Chinese species, T. striatus differs from the type species by the clearly presented discal spots and whitish lines along veins on the forewing (Fig. 5). The male genitalia are also very similar, but gnathos is narrower and longer, and the distal part of valva is proportionally shorter. Female unknown. Male genitalia (Figs. 16, 16a, b). Described and illustrated by Gozmány (1978, Taf. 36, Fig. 80). Material examined. 4♂, Chiang Mai, Doi Pui, 1300 m, 1–4 IX 1987 (Moriuti, Saito, Arita, Yoshimoto), gen. slide no. 5828/Park. Distribution. Nepal, Thailand (new record; Chiang Mai).
Genus Tegenocharis Gozmány, 1973 Tegenocharis Gozmány, 1973, Khumbu Himal. 4: 429; 1978: 135: Wu, 1997: 199. Type species: Tegenocharis tenebrans Gozmány, 1973.
Genus Eurodachtha Gozmány, 1978 Microlep. Palaearctica 5: 151. Type species: Lecithocera pallicornella Staudinger, 1859. TL: Spain. Eurodachtha Gozmány has narrow, elongated forewing, with all veins: R3 free from R4 + 5, M2 and M3 free; CuA1 and CuA2 far from each other; hindwing with M2 atrophied, often with only vestige (originally described as M2 and M3 coincident); M3 and CuA1 short
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Figs. 12–15. Male genitalia, abdominal segments, and female genitalia. 12, Procharista ranongensis sp. nov. sl. No. 5827; 12a, ditto, aedeagus; 12b, ditto, abdominal segment; 13, Procharista spectabilosa sp. nov., sl. No. 5855; 13a, ditto, aedeagus; 13b, ditto, abdominal segment; 13c, ditto, enlargement of gnathos; 14, Urolaguna heosa Wu, sl. No. 5839; 14a, ditto, aedeagus; 14b, ditto, abdominal segment; 15. Female genitalia of Urolaguna heosa Wu, sl. No. 5849.
stalked or connate; CuA2 arising from middle or beyond (Fig. 11). Abdominal tergites have sparsely spiniform setae which are easily removable. Five species were known only in the Mediterranean subregion, but no species has been known from the Oriental Region. The following new species is reported for the first time from the Oriental Region. Eurodachtha rotundina sp. nov. (Figs. 6, 6a, 6b, 11, 17, 17a, b) Type. Holotype. ♂, Thailand, Nakhon Noyak, Khao Yai, 800 m, 20 VI 1983 (Kuroko, Moriuti, Arita, Yoshiyasu), gen. slide no. 5829/Park, wings prep. no. 5851/Park.
Diagnosis. The species is more or less similar to E. nigralba Gozmány, which was described from Syria, but differs from the latter by the yellowish brown forewing and the different shape of the valva of the male genitalia, with strongly bent cucullus. Description. Wingspan, 12.5 mm. Head yellowish brown, with grayish orange erect scales laterally. Scape of antenna elongated, slightly broadened toward apex, grayish orange dorsally, orange white ventrally; flagellum yellowish brown, with indistinct dark brown annulations; apex pale orange gray. Labial palpus slender, strongly upturned; second segment slender, shiny orange white on both surfaces; 3rd segment as long as 2nd, dark brown ventrally. Tegula and thorax yellowish brown. Forewing elongate, lanceolate, with acute
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Figs. 16–19. Male genitalia, abdominal segments, and female genitalia. 16, Tegenocharis tenebrans Gozmány, sl. No. 5828; 16a, ditto, aedeagus; 16b, ditto, abdominal segment; 17, Eurodachtha rotundina sp. nov., sl. No. 5829; 17a, ditto, aedeagus; 17b, ditto, abdominal segment; 18, Trichoboscis pansarista Meyrick, sl. No. 5840; 18a, ditto, aedeagus; 18b, enlargement of aedeagus, sl. No. 5087; 19, Female genitalia of Trichoboscis pansarista Meyrick, sl. No. 5857.
apex; ground color uniform metallic yellowish brown, without any markings; costa nearly straight prior to 4/5 length; termen strongly oblique; fringe concolorous; venation with R1 arising before middle of cell; R2 arising nearly from middle; R3 free from R4 + 5; R4 and R5 stalked near middle; R5 terminated to apex; M1 far from R4 + 5 at base; M2 nearly parallel to M1; M2 and M3 free; CuA1 and CuA2 nearly connate, proportionally short; cell closed; apex acute; termen extremely oblique. Hindwing pale orange gray; costa nearly straight; apex acute; termen oblique, slightly concave medially; venation with
Rs and M1 stalked at near middle; M2 atrophied, with only vestige; M3 and CuA1 connate; CuA2 arising from middle of cell. Hind tibia with pale grayish orange hair-like scales dorsally; spurs proportionally long, longer one about 1/2 length of hind tibia; tarsi concolorous. Abdominal tergites have sparsely spiniform setae dorsally. Female unknown. Male genitalia (Figs. 17, 17a, b). Uncus-lobes short, semiovate. Gnathos short, prorptionally small. Costal bar sharply angled at middle. Valva broad at basal 1/3, with large cavity on surface; distal 2/ 5 nearly parallel, curved distally, dense setosed, with rounded apex;
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costa strongly concaved, with dense setae on surface; sacculus broad, with long setae. Juxta shield-shaped; caudal margin concave with narrowly produced laterocaudal processes. Aedeagus stout, as long as valva, with various length of short strong spines beyond middle. Distribution. Thailand (Nakhon Nayok). Genus Trichoboscis Meyrick, 1929 Exot. Microlep. 3: 526; Gozmány, 1978: 176. Wu, 1997: 198. Type species: Trichoboscis pansarista Meyrick, 1929. Trichoboscis Meyrick is easily distinguished from its allies by the antenna with a notch at basal 1/5 and the unique shape of the labial palpus of the male: the 2nd segment with a long scales-tuft dorsally and the 3rd segment short, stout. The labial palpus of the female is normally recurved. Abdominal tergites have spinous zones dorsally. Two species are known from India and China. Trichoboscis pansarista Meyrick, 1929 (Figs. 7, 7a, 7b, 18, 18a, b, 19) Trichoboscis pansarista Meyrick, 1929, Exot. Microlep. 3: 526; Clarke, 1965: 255; Gozmány, 1978: 176. TL: Sikkim, India. Type in the Natural History Museum (BMNH), London. Diagnosis. Forewing length 15 mm. The male genitalia of the species are very similar those of T. (Clarke, 1965, Pl. 61, Figs. 2–2b), but can be distinguished by the less slender valva and more thorn-like spines in the aedeagus. The female genitalia also differs by a single plate of signum in T. pansarista Meyrick, instead of three plate of signa in T. crocosema Meyrick. Male genitalia (Figs. 18, 18a, b). Illustrated by Clarke (1965, Pl. 127, Figs. 1c–d), Gozmány (1978, Taf. 43, Fig. 106). Female genitalia (Fig. 19). Eighth abdominal segment weakly sclerotized, slightly emarginated at middle on caudal margin. Apophyses anteriores about 1/2 length of apophyses posteriors. Antrum memberanous. Ductus bursae short, funnel-like, about 2/3 length of corpus bursae. Corpus bursae ovate, bearing numerous, very short corn-shaped spines with broad basal part on internal surface; signum small, semiovate, and radiated. Material examined. 1♂, Vietnam, Cuc Phuong Nat. Park, 300 m, 2 V 2005 (KT Park and SR Kim), gen. slide no. CIS-5087/Park; 1♂, same locality as above, 30 IV–1V 2005 (KT Park and SR Kim); 7♂, Vietnam, Bac Khan Prov., Ba Be National Park, 240 m, 26–28 VII 2006 (Park, Chae, and Cuong); 1♂, same locality as above, 21–26 Jun 2008 (JB Heppner); 2♂, Thailand, Nakhon Nayok, Khao Yai, 800 m, 21–22 VI 1983 (Kuroko, Moriuti, Saito, Arita, Yoshiyasu); 1♂, 1♀, Thailand, Chiang Mai, Khao Soi Dao, 480 m, 24–25 VIII 1987 (Moriuti, Saito, Arita, Yoshimoto), gen. slide no. 5840/Park(♂) and 5857/Park(♀); 1♂, same locality, 22 IX 1987 (Moriuti, Saito, Arita, Yoshiyasu).
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Distribution. India (North), Thailand (new record; North and Central), Vietnam (new record; North). Acknowledgments Authors are grateful to T. Hirowatari, Osaka Prefecture University, Osaka, Japan, for the loan of valuable specimens for this study, and M Thomas, Florida State Collection of Arthropods, Division of Plant Industry, FDACS, Gainesville, FL, USA, for his allowance in using all available equipments, including Automontage microscope. A part of the financial support was given by the National Institute of Biological Resource, Korea. References Clarke, J.F.G., 1965. Catalogue of the type specimens of Microlepidoptera in the British Museum (Natural History) described by Edward Meyrick. Vol. 5, Vol. 5. London, 581 pp. Gozmány, L., 1973. Symmocid and Lecithocerid moths (Lepidoptera) from Nepal. Khumbu Himalaya 4 (3), 413–444. Gozmány, L., 1978. Lecithoceridae. In: Amsel, H.G., Gregor, F., Reisser, H. (Eds.), Microlepidoptera Palaeartica, Vol. 5. Georg Fromme and Co., Wien. 306 pp. Kornerup, A., Wanscher, J.H., 1983. Methuen Handbook of Colour. 3rd ed. 252 pp. Methuen and Co. Ltd., London. Meyrick, E., 1922. New Micro-Lepidoptera. Zool. Meded. 7, 80–89. Meyrick, E. 1929. Exotic Microlepidoptera 3: 521–526. Marloborough, Wilts. Park, K.T., 2001. A new species of the genus Tisis (Lepidoptera: Lecithoceridae) from Thailand. Korean J. Syst. Zool. 17 (1), 59–63. Park, K.T., 2002a. Taxonomic review of the genus Torodora Meyrick in Thailand, with descriptions of fifteen new species (Lepidoptera: Lecithoceridae). Insecta Koreana 19 (2), 147–166. Park, K.T., 2002b. Four new species of the genus Antiochtha Meyrick (Lepidoptera: Lecithoceridae) from Thailand. J. Asia-Pacific Entomol. 5 (2), 139–143. Park, K.T., 2003a. Two new species of the genus Tisis from Thailand and notes on Tisis elegans (Insecta: Lepidoptera: Lecithoceridae). Species Divers. 8, 267–274. Park, K.T., 2003b. Three new species of the genus Hygroplasta Meyrick (Lepidoptera, Lecithoceridae) from the Philippines and Thailand. J. Asia-Pacific Entomol. 6 (2), 131–135. Park, K.T., 2003c. Thubana species (Lepidoptera, Lecithoceridae) in Thailand, with descriptions of twelve new species. J. Asia-Pacific Entomol. 6 (2), 137–150. Park, K.T., 2004. Genus Homaloxestis Meyrick of Thailand, with description of five new species (Lepidoptera, Lecithoceridae). J. Asia-Pacific Entomol. 7 (1), 33–43. Park, K.T., 2005a. Four new species of the genus Frisilia Walker from Thailand (Lepidoptera: Lecithoceridae, Lecithocerinae). Tinea 18 (Suppl. 3), 55–62. Park, K.T., 2005b. Genus Lecithocera Herrich-Shäffer of Thailand (I): descriptions of three new species (Lepidoptera, Lecithoceridae). J Asia-Pacific Entomo. 8 (3), 233–237. Park, K.T., 2006a. Genus Lecithocera of Thailand (II): descriptions of four new species (Lepidoptera, Lecithoceridae). Tinea 19 (2), 98–103. Park, K.T., 2006b. Genus Lecithocera of Thailand (III): new records of five species of the genus (Lepidoptera, Lecithoceridae). J. Asia-Pacific Entomol. 9 (4), 313–316. Park, K.T., 2007. Three new species of Torodora Meyrick from Thailand (Lepidoptera, Lecithoceridae). Shilap Revista Lepidop. 35, 23–28. Wu, C., 1994. The Lecithoceridae (Lepidoptera) of China, with description of new taxa. Sinzoologica 11, 123–154. Wu, C., 1997. Lepidoptera, Lecithoceridae. Fauna Sinica, Insecta., Vol. 7. Science Press, Beijing, pp. 1–302.