Nomenclature for factors of the HLA system, update October 2015

Human Immunology 77 (2016) 440–444

Contents lists available at ScienceDirect

www.ashi-hla.org

journal homepage: www.elsevier.com/locate/humimm

Nomenclature for factors of the HLA system, update October 2015 Compiled by Steven G.E. Marsh, for the WHO Nomenclature Committee for Factors of the HLA System

The following sequences have been submitted to the Nomenclature Committee since the September 2015 nomenclature update and, following agreed policy, have been assigned official allele designations [1]. Full details of all sequences will be published in a forthcoming report. Below are listed the newly assigned sequences (Table 1) and confirmations of previously reported sequences (Table 2). The accession number of each sequence is given and these can be used to retrieve the sequence files from the EMBL, GenBank or DDBJ data libraries. Although accession numbers have been assigned by the data-libraries and most sequences are already available, there is

still the possibility that an author may not yet have allowed the sequence to be released; in such a case you will have to contact the submitting author directly. Additional information pertaining to new sequences is often included in the publications describing these alleles; a listing of recent publications that describe new HLA sequences is given in Table 3. All new and confirmatory sequences should now be submitted directly to the WHO Nomenclature Committee for Factors of the HLA System via the IMGT/HLA Database using the sequence submission tool provided [2]. The IMGT/HLA Database may be accessed via the World Wide Web at: www.ebi.ac.uk/ipd/imgt/hla.

Table 1 New sequences. HLA allele

Cell identification

Accession number

Submitting author

A⁄01:197 A⁄01:198 A⁄02:01:121 A⁄02:599 A⁄02:600 A⁄02:601 A⁄03:01:01:06 A⁄03:154:02 A⁄03:235 A⁄03:236 A⁄24:324 A⁄24:325 A⁄24:326 A⁄30:01:11 A⁄31:104 A⁄32:80 A⁄33:105 A⁄68:138 A⁄68:139 A⁄74:27 B⁄07:257 B⁄07:258 B⁄07:259 B⁄07:260 B⁄08:157 B⁄08:158

HG00010818 HG00010821 NT01279 HG00010827 HG00010828 HG00010829 NT01275 15207482 HG00010822 HG00010823 HG00010816 SZGSQ24 275881 NT01272 HG00010808 HG00010824 HG00010809 HG00010812 NT01278 HG00010825 HG00010700 HG00010702 HG00010857 HG00010858 HG00010836 HG00010838

KT710866 KT600076 KT819313 KT600073 KT600078 KT710868 KT819308 KT803854 KT760491 KT600070 KT600071 KT826594 KT878496 KT819305 KT760485 KT760482 KT760493 KT710867 KT819312 KT760483 KT438907 KT438910 KT760505 KT760506 KT600096 KT710870

Histogenetics, Ossining, USA Histogenetics, Ossining, USA Carolyn K Hurley, Washington Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Carolyn K Hurley, Washington Daniel Fuerst, Ulm, Germany Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Su-Qing Gao, Shenzhen, China Elena Longhi, Milano, Italy Carolyn K Hurley, Washington Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Carolyn K Hurley, Washington Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA

DC, USA

DC, USA

DC, USA

DC, USA

⇑ Addrress for correspondence: Professor Steven G.E. Marsh, Anthony Nolan Research Institute, Royal Free Hospital, Pond Street, London NW3 2QG, United Kingdom. E-mail address: [email protected] URLs: http://hla.alleles.org, http://www.anthonynolan.org/hig, http://www.ebi.ac.uk/ipd/imgt/hla http://dx.doi.org/10.1016/j.humimm.2016.01.021 0198-8859/Ó 2016 American Society for Histocompatibility and Immunogenetics. Published by Elsevier Inc. All rights reserved.

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S.G.E. Marsh / Human Immunology 77 (2016) 440–444 Table 1 (continued) HLA allele ⁄

B 13:86 B⁄13:87 B⁄14:51 B⁄15:04:03 B⁄15:375N B⁄15:376 B⁄15:377Q B⁄18:01:24 B⁄27:146 B⁄35:301 B⁄35:302 B⁄35:303 B⁄35:304 B⁄35:305 B⁄35:306 B⁄38:59 B⁄39:108 B⁄39:109 B⁄39:110 B⁄40:318 B⁄40:319 B⁄40:320 B⁄40:321 B⁄41:42 B⁄41:43 B⁄41:44 B⁄42:22 B⁄44:03:31 B⁄44:03:32 B⁄44:128:02 B⁄44:235 B⁄45:18 B⁄48:38 B⁄51:196 B⁄51:197 B⁄51:198 B⁄56:50 B⁄57:81 B⁄57:82 B⁄58:59:02 B⁄58:74 B⁄58:75 B⁄58:76 C⁄01:115 C⁄01:116 C⁄01:117N C⁄02:06:02 C⁄02:10:01:02 C⁄02:109 C⁄03:04:49 C⁄03:57:02 C⁄03:304 C⁄03:305 C⁄03:306 C⁄03:307 C⁄04:218 C⁄04:219 C⁄04:220 C⁄04:221 C⁄04:222 C⁄04:223 C⁄04:224 C⁄04:225N C⁄04:226 C⁄05:125 C⁄05:126 C⁄06:167 C⁄06:168 C⁄06:169 C⁄06:170 C⁄07:257:04 C⁄07:460 C⁄07:461 C⁄07:462 C⁄07:463

Cell identification

Accession number

Submitting author

HG00010840 BY01270 HG00010856 HG00010849 60750978 HG00010852 NT01277 TMO27647 HG00010830 HG00010839 HG00010843 HG00010845 HG00010846 96083726 93103407 HG00010706 HG00010704 HG00010705 HG00010708 HG00010709 HG00010831 HG00010834 BY01275 HG00010833 15700134 NT01281 HG00010701 HG00010861 NT01280 LUMC-B-126 HG00010841 HG00010832 HG00010860 HG00010853 HG00010854 HG00010855 282336 HG00010850 BY01274 HG00010848 HG00010699 HG00010847 NT01271 HG00010862 81136719 NT01273 BY01271 NT01276 HG00010889 LUMC-C-117 HG00010737 HG00010736 HG00010738 HG00010739 HG00010894 HG00010741, HG00010742 HG00010743 HG00010865 HG00010898 HG00010899 HG00010900 HG00010901 HG00010902 LUMC-C-118 HG00010724 HG00010886 HG00010711 HG00010712 HG00010864 HG00010866 HG00010870 HG00010716 HG00010717 HG00010718 HG00010719

KT600089 KT764976 KT600097 KT600091 KT768091 KT600094 KT819311 KT807796 KT600085 KT600081 KT760498 KT760503 KT760495 KT803855 KT803856 KT355342 KT438911 KT315593 KT315595 KT438909 KT600082 KT600092 KT819316 KT760494 KT852371 KT819315 KT282157 KT600093 KT819314 LN887899 KT760501 KT760496 KT600090 KT760507 KT710873 KT600084 KT821556 KT600083 KT764975 KT760500 KT315597 KT710871 KT764979 KT760521 KT803858 KT819306 KT764974 KT819309 KT600113 LN887896 KT282169 KT282176 KT361896 KT282168 KT710876 KT282167, KT282174 KT235658 KT760516 KT760514 KT600112 KT600123 KT600106 KT760508 LN887897 KT361897 KT600122 KT315601 KT235659 KT600116 KT710874 KT600119 KT355373 KT355367 KT315604 KT282172

Histogenetics, Ossining, USA Carolyn K Hurley, Washington DC, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Antonio Balas, Madrid, Spain Histogenetics, Ossining, USA Carolyn K Hurley, Washington DC, USA Antonio Balas, Madrid, Spain Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Daniel Fuerst, Ulm, Germany Daniel Fuerst, Ulm, Germany Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Carolyn K Hurley, Washington DC, USA Histogenetics, Ossining, USA Antonio Balas, Madrid, Spain Carolyn K Hurley, Washington DC, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Carolyn K Hurley, Washington DC, USA Jacqy DH Anholts, Leiden, The Netherlands Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Elena Longhi, Milano, Italy Histogenetics, Ossining, USA Carolyn K Hurley, Washington DC, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Carolyn K Hurley, Washington DC, USA Histogenetics, Ossining, USA Daniel Fuerst, Ulm, Germany Carolyn K Hurley, Washington DC, USA Carolyn K Hurley, Washington DC, USA Carolyn K Hurley, Washington DC, USA Histogenetics, Ossining, USA Jacqy DH Anholts, Leiden, The Netherlands Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Jacqy DH Anholts, Leiden, The Netherlands Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA (continued on next page)

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S.G.E. Marsh / Human Immunology 77 (2016) 440–444

Table 1 (continued) HLA allele

Cell identification

Accession number

Submitting author

C⁄07:464 C⁄07:465 C⁄07:466 C⁄07:467 C⁄07:468 C⁄07:469 C⁄07:470 C⁄07:471 C⁄07:472 C⁄07:473 C⁄07:474 C⁄07:475 C⁄07:476N C⁄08:126 C⁄08:127N C⁄12:168 C⁄12:169 C⁄12:170 C⁄12:171 C⁄14:73 C⁄15:118 C⁄15:119 C⁄15:120 C⁄16:88 C⁄16:89N C⁄18:10 DRB1⁄01:65:02 DRB1⁄03:126 DRB1⁄03:127 DRB1⁄07:67 DRB1⁄08:78N DRB1⁄11:02:05 DRB1⁄13:211 DRB1⁄13:212 DRB1⁄14:169 DRB1⁄14:170 DRB1⁄15:01:28 DRB1⁄15:128 DRB1⁄15:129N DRB3⁄01:18 DRB3⁄01:19 DRB3⁄02:02:06 DRB3⁄02:30 DRB3⁄02:31 DRB3⁄02:32 DQA1⁄01:13 DQB1⁄02:65 DQB1⁄03:216 DQB1⁄03:217 DQB1⁄03:218 DQB1⁄04:33 DQB1⁄05:66:02 DQB1⁄05:89:02 DQB1⁄05:110N DQB1⁄05:111 DQB1⁄06:198 DPA1⁄02:01:01:02 DPB1⁄14:01:03 DPB1⁄498:01 DPB1⁄499:01 DPB1⁄500:01 DPB1⁄501:01 DPB1⁄502:01 DPB1⁄503:01 DPB1⁄504:01 DPB1⁄505:01 DPB1⁄506:01 DPB1⁄507:01N DPB1⁄508:01 DPB1⁄509:01 DPB1⁄510:01 DPB1⁄511:01 DPB1⁄512:01 DPB1⁄513:01 DPB1⁄514:01

HG00010721 HG00010723 HG00010868 HG00010869 HG00010871 HG00010872 HG00010875 HG00010876 HG00010878 HG00010879 HG00010880 HG00010882 HG00010722 HG00010725 13QB5759 HG00010728 HG00010729 HG00010888 281512 HG00010897 HG00010731, HG00010732 HG00010890 HG00010892 HG00010727 HG00010730 NT01274 HG00010916 HG00010776 HG00010912 HG00010915 HG00010917 HG00010922 HG00010919 HG00010920 HG00010780 HG00010923 HG00010914 HG00010913 LUMC-DRB1-124 HG00010937 HG00010938 LUMC-DRB3-122 HG00010939 HG00010940 HG00010941 60753693 SNUH150818_27385852 HG00010903 HG00010905 93023157 HG00010910 IU060320150001 HG00010906 HG00010908 HG00010909 HG00010911 BY01256 HG00010931 UNC11 HG00010789 HG00010791 HG00010793 HG00010794 HG00010796 HG00010797 HG00010798 HG00010800 HG00010802, HG00010803 HG00010804 HG00010806 HG00010925 HG00010926 HG00010928 HG00010930 HG00010932, HG00010933

KT315606 KT282179 KT760522 KT760512 KT600103 KT760510 KT600111 KT760524 KT600120 KT760520 KT760532 KT760528 KT315600 KT361894 KP760115 KT282164 KT355374 KT600105 KT878495 KT760529 KT282163, KT282166 KT760519 KT760530 KT282178 KT282159 KT819307 KT600135 KT282190 KT760538 KT600137 KT600130 KT760539 KT760542 KT760540 KT438916 KT760543 KT760541 KT600131 LN887900 KT781582 KT781581 LN887898 KT781583 KT781584 KT781585 KT807797 KT878020 KT710877 KT760536 KT803857 KT600129 KT728007 KT760533 KT600125 KT760534 KT600126 KT716046 KT760552 KT001470 KT315617 KT315616 KT355357 KT438920 KT361915 KT361912 KT282196 KT235669 KT438918, KT361911 KT235664 KT315618 KT600138 KT600145 KT760547 KT600142 KT760546, KT600143

Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Janette Kwok, Pokfulam Road, Hong Kong Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Elena Longhi, Milano, Italy Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Carolyn K Hurley, Washington DC, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Jacqy DH Anholts, Leiden, The Netherlands Histogenetics, Ossining, USA Histogenetics, Ossining, USA Jacqy DH Anholts, Leiden, The Netherlands Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Antonio Balas, Madrid, Spain Oh-Joong Kwon, Seoul, South Korea Histogenetics, Ossining, USA Histogenetics, Ossining, USA Daniel Fuerst, Ulm, Germany Histogenetics, Ossining, USA Jeremy Sherrill, Indianapolis, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Carolyn K Hurley, Washington DC, USA Histogenetics, Ossining, USA Eric Weimer, Chapel Hill, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA

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S.G.E. Marsh / Human Immunology 77 (2016) 440–444 Table 1 (continued) HLA allele

Cell identification

Accession number

Submitting author

DPB1⁄515:01 DPB1⁄516:01 DPB1⁄517:01

HG00010934 HG00010935 HG00010936

KT760544 KT760553 KT760549

Histogenetics, Ossining, USA Histogenetics, Ossining, USA Histogenetics, Ossining, USA

Table 2 Confirmatory sequences. HLA allele

Cell identification

Accession number

Submitting author

A⁄01:190 A⁄11:190 B⁄07:02:48 B⁄18:01:21 B⁄38:01:08 B⁄40:01:40 C⁄02:91 C⁄04:01:01:06 C⁄05:111 C⁄07:02:49 C⁄14:70 C⁄16:01:18 DQB1⁄02:53Q DQB1⁄03:200 DQB1⁄06:04:01 DPA1⁄02:02:02

281368 BY01273 HG00010859 BY01272 HG00010707 BY01214 HG00010733 BY01272 HG00010884 BY00761 HG00010896 NT01276 81135639 HG00010904 NT01262, NT01263 BY01262, BY01263

KT878494 KT764980 KT600086 KT764977 KT315592 KJ623754 KT282165 KT764978 KT600102 HQ700658 KT760517 KT819310 KT803859 KT760535 KT591471 KT716050

Elena Longhi, Milano, Italy Carolyn K Hurley, Washington Histogenetics, Ossining, USA Carolyn K Hurley, Washington Histogenetics, Ossining, USA Carolyn K Hurley, Washington Histogenetics, Ossining, USA Carolyn K Hurley, Washington Histogenetics, Ossining, USA Carolyn K Hurley, Washington Histogenetics, Ossining, USA Carolyn K Hurley, Washington Daniel Fuerst, Ulm, Germany Histogenetics, Ossining, USA Carolyn K Hurley, Washington Carolyn K Hurley, Washington

Table 3 Recently published sequences.

DC, USA DC, USA DC, USA DC, USA DC, USA DC, USA

DC, USA DC, USA

Table 3 (continued) HLA allele

HLA allele

References

A⁄02:07:07 A⁄02:48 A⁄02:251 A⁄24:313 A⁄29:01:08 A⁄29:01:09 A⁄33:95 B⁄35:270 B⁄35:273 B⁄46:01:20 C⁄08:01:09 DRB1⁄03:04:01 DRB1⁄03:07 DRB1⁄03:15 DRB1⁄04:03:02 DRB1⁄04:14 DRB1⁄04:15 DRB1⁄04:16 DRB1⁄04:25 DRB1⁄04:26 DRB1⁄04:31 DRB1⁄04:32 DRB1⁄07:04 DRB1⁄08:04:01 DRB1⁄08:06 DRB1⁄08:07 DRB1⁄08:10 DRB1⁄08:11 DRB1⁄08:14 DRB1⁄08:16 DRB1⁄08:35 DRB1⁄08:77 DRB1⁄11:05 DRB1⁄11:06:01 DRB1⁄11:08:02 DRB1⁄11:09 DRB1⁄11:11:01 DRB1⁄11:13:02 DRB1⁄11:14:01 DRB1⁄11:19:01

[3] [4] [4] [5] [6] [7] [8] [9] [9] [10] [11] [12] [12] [12] [12] [12] [12] [12] [12] [12] [12] [12] [12] [12] [12] [12] [12] [12] [12] [12] [12] [12] [12] [12] [12] [12] [12] [12] [12] [12]

⁄

DRB1 11:21 DRB1⁄11:29:01 DRB1⁄13:03:01 DRB1⁄13:04 DRB1⁄13:06 DRB1⁄13:08 DRB1⁄13:10 DRB1⁄13:11:01 DRB1⁄13:14:02 DRB1⁄13:17 DRB1⁄13:20 DRB1⁄13:26:01 DRB1⁄13:36 DRB1⁄14:02:01 DRB1⁄14:06:01 DRB1⁄14:08 DRB1⁄14:10 DRB1⁄14:14 DRB1⁄14:17 DRB1⁄14:21 DRB1⁄14:24 DRB1⁄15:02:02 DRB1⁄15:07:01 DRB1⁄16:04 DRB1⁄16:07 DRB1⁄16:36

References [12] [12] [12] [12] [12] [12] [12] [12] [12] [12] [12] [12] [12] [12] [12] [12] [12] [12] [12] [12] [12] [12] [12] [12] [12] [13]

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